CDS , the red circle in the middle of the nest is the spot where I would focus, some very old places are still trapped and the sentinels were probably just as scared as anyone else about digging them up, but they still marked them out or "freshened" them up. Some are probably marked out by other treasure hunters who came and went as well. Also the all seeing eye is often looking right at the spot, I have seen many of them and they all look towards it or watch over it , they do not lie and it is the eye of God. They are sometimes hard to see, but they didn't want it to be easily found.
Sometimes they watch over where it is located deep in the ground, but not where you would dig to get at it, the entrance may be several yards away. Occasional they are just looking at a marker or a site as a whole, but most often they look at a marker or something important when the eye is part of a face or animal, etc. When the eye stands alone as an "eye" it is watching over a treasure or treasure site and it is protecting it with the magic they believed in..
Tinhorn, there were ancient groups looting out ancient vaults and crypts, it depends on what you consider ancient. It's such a messed up timeline of "who did what" is the reason I say to focus more on the "Where".